# Selenium nanoparticles and wheat straw biochar synergistically alleviate combined drought–heat stress in wheat (Triticum aestivum L.) by modulating antioxidant defense, photosynthetic efficiency, and ion homeostasis
> Firdous H.

URL kanonis: https://discover.unhas.ac.id/publications/pub_scopus_105044174288
Jurnal / Konferensi: Plant Signaling and Behavior
Tahun terbit: 2026
DOI: https://doi.org/10.1080/15592324.2026.2681236
ISSN: 15592316
Kuartil SJR: Q1
Citations: 0

## Authors
- Firdous H.

## Abstract
) in a completely randomized design with three replications. Combined DS+HS severely impaired growth, photosynthetic function, and grain yield, while markedly elevating oxidative damage markers and disrupting ionic homeostasis. Notably, combined Se NPs + biochar application produced superior synergistic amelioration relative to either amendment alone, restoring net photosynthetic rate and grain yield to approximately 89%-90% and 88% of control values, respectively, alongside significant reductions in oxidative stress indicators. Gene expression analysis confirmed marked upregulation of key stress-responsive genes (TaSOD1, TaDREB2, TaHSP70, TaNHX1) and coordinated antioxidant enzyme activation. These findings establish that soil co-application of Se NPs and biochar is an effective, agronomically feasible strategy to sustain wheat productivity under the increasingly prevalent combined DS+HS conditions in Pakistan's irrigated cropping systems.

## Keywords
- Biochar
- Photosynthesis
- Antioxidant
- Oxidative stress
- Agronomy
- Biology
- Reactive oxygen species
- Selenium
- Amendment
- Straw
- Food science
- Glutathione
- Photosynthetic efficiency
- Chemistry
- Poaceae
- Crop yield
- Genetically modified crops
- Botany
- Lipid peroxidation
- Biomass (ecology)
- Stomatal conductance
- Biotechnology
- Wheat grain
- Grain yield
- Biochemistry

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Sumber: Discover Unhas — RIMS Universitas Hasanuddin.
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